This is a list of Richard D. Alexander’s writings, arranged first by category and then by year. Most of these, including all of his professional scientific articles, are available for download (indicated by “PDF” to the left of the entry). RDA had a broad conception of a “professional” work, including for instance some letters to newspapers, entries in society newsletters or encyclopedias, and administrative reports. This page follows his own conventions, with all of the writings listed here as professional having been designated by RDA specifically as such.

Either scroll down or click on one of these categories to browse, or else use the Find command (Ctrl-F or Cmd-F) to search this page.

RDA’s poetry is distributed among various works, so individual poems can be difficult to locate. Click here to access the gradually accumulating alphabetical collection of Richard D. Alexander’s poetry.

Professional Books & Monographs
Professional Articles
Unpublished Manuscripts
Nonprofessional Works

Professional Books & Monographs


[PDF] 1956. A Comparative Study of Sound Production in Insects, With Special Reference to the Singing Orthoptera and Cicadidae of the Eastern United States. Ph.D. Dissertation: The Ohio State University, xviii + 529 pp.  [The abstract, as published in Dissertation Abstracts, is here.]


[PDF] 1967. Singing Insects: Four Case Histories in the Study of Animal Species. Rand McNally Pattern of Life Series. Chicago: Rand McNally, 86 pp.


1979. Darwinism and Human Affairs, University of Washington Press, Seattle. xiv + 317 pp.
[Available to purchase from Amazon here and from AbeBooks here].


1983. The Australian Crickets: (Orthoptera: Gryllidae). 376 new species, 40 new genera, monograph 22, Philadelphia Academy of Natural Sciences, Philadelphia, 477 pp. (D. Otte and R. D. Alexander).
[Available to purchase from the Academy of Natural Sciences here, and to view or purchase on Google here].


[PDF] 1987. The Biology of Moral Systems, xxii + 303 pp., Hawthorne, N. Y.: Aldine de Gruyter.
[Available to purchase from Amazon here and from Google here].




[PDF] 1981. Natural Selection and Social Behavior: Recent Research and New Theory, New York: Chiron Press, ca. 500 pp. (R. D. Alexander and D. W. Tinkle, eds.).


1991. The Biology of the Naked Mole Rat, Princeton Univ. Press. (P. W. Sherman, J. Jarvis, and R. D. Alexander, eds.).
[Available to purchase from Amazon here and Princeton here].



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Professional Articles


[PDF] 1955. Capturing the melodies of insect singers. Magnetic Film and Tape Recording Magazine 2(5A): 34-37. [The entire issue is included in this file].

[PDF] 1957. Sound production and associated behavior in insects. Ohio Journal of Science 57(2): 101-113.

[PDF] 1957. The taxonomy of the field crickets of the eastern United States(Orthoptera: Gryllidae: Acheta). Annals of the Entomological Society of America 50(6): 584-602.
Reprinted in P. Barbosa and T. M.. Peters (eds.), Readings in Entomology, W. E.. Saunders Co., 1972.

[PDF] 1957. Nemobius melodius, a new species of cricket from Ohio (Orthoptera: Gryllidae). Ohio Journal of Science 57(3): 148-152. (E. S. Thomas and R.D. Alexander).

[PDF] 1957. The song relationships of four species of ground crickets (Orthoptera: Gryllidae: Nemobius). Ohio Journal of Science 57(3): 153-163.

[PDF] 1957. The periodical cicadas. Wheaton Club Bulletin 2(2): 23-26.

[PDF] 1958. Studies on the acoustical behavior of seventeen-year cicadas (Homoptera: Cicadidae: Magicicada). Ohio Journal of Science 58(2): 107-127. (R.D. Alexander and T. E. Moore).

[PDF] 1958. The periodical cicada complex. Proceedings of the Tenth International Congress of Entomology 1: 349-356 (T. E. Moore and R. D. Alexander).

[PDF] 1959. The courtship and copulation of Pasimachus punctulatus (Haldemann) (Coleoptera: Carabidae). Annals of the Entomological Society of America 52(4): 485.

[PDF] 1959. Systematic and behavioral studies on the crickets of the Nemobius fasciatus group (Orthoptera: Gryllidae: Nemobiinae). Annals of the Entomological Society of America 52(5): 591-605. (R. D. Alexander and E. S. Thomas).


[PDF] 1960. Communicative mandible-snapping in Acrididae (Orthoptera). Science 132 (3240): 152-153.

[PDF] 1960. Sound communication in Orthoptera and Cicadidae. In: Lanyon WE, Tavolga WN, editors. Animal Sounds and Communication. Washington D.C.: American Institute of Biological Sciences, p 38-92. [Original version included 5 sets of songs on a phonograph record.]

[PDF] 1960. Allochronic speciation in field crickets, and a new species, Acheta veletis. Evolution 14(3): 334-336. (R.D. Alexander and R. Bigelow).
Reprinted in Papers on Evolution, Ehrlich, Holm and Raven (eds.) 1969. Little, Brown Publ.

[PDF] 1961. Aggressiveness, territoriality, and sexual behavior in field crickets (Orthoptera: Gryllidae). Behaviour 17: 130-223, 63 figs., 31 tab.
Reprinted (in part) in Territoriality: Landmark Papers in Animal Behavior, A. W.. Stokes (ed.), Dowden, Hutchinson, and Ross, Pub. 1974, p. 364.

[PDF] 1962. Two introduced field crickets new to eastern United States (Orthoptera: Gryllidae).Annals of the Entomological Society of America 55(1): 90-94 (R.D. Alexander and T. J.. Walker).

[PDF] 1962. The role of behavioral study in cricket classification. Systematic Zoology 11(2): 53-72.

[PDF] 1962. The evolutionary relationships of 17-year and 13-year cicadas, and three new species (Homoptera: Cicadidae: Magicicada). Miscellaneous Publications of the University of Michigan Museum of Zoology 121: 1- 59. (R. D. Alexander and T. E. Moore).

[PDF] 1962. Invertebrate bio-acoustics: 1961. Bio-Acoustics Bulletin 2(2 & 3): 7-10.

[PDF] 1962. Evolutionary change in cricket acoustical communication. Evolution 16(4): 443-467.

[PDF] 1962. Systematic and behavioral studies on the meadow grasshoppers of the Orchelimum concinnum group (Orthoptera: Tettigoniidae). Occasional Papers of the University of Michigan Museum of Zoology 626: 1-31. (E.S. Thomas and R.D. Alexander).

[PDF] 1963. The evolutionary differentiation of stridulatory signals in beetles (Insecta: Coleoptera). Animal Behaviour 11(1): 115. (R. D. Alexander, T. E. Moore, and R. E.. Woodruff).

[PDF] 1963. Sub-social behavior in a burrowing cricket, Anurogryllus muticus (De Geer) (Orthoptera: Gryllidae). Ohio Journal of Science 63(1): 19-24 (M. J. West and R.D. Alexander).

[PDF] 1963. Mating behavior and the origin of insect wings. Occasional Papers of the University of Michigan Museum of Zoology 628: 1-19. (R.D. Alexander and W. L. Brown).

[PDF] 1963. Cricket. Encyclopaedia Britannica. Vol. 6:753.

[PDF] 1963. Invertebrate bio-acoustics: 1962. Bio-Acoustics Bulletin 3(3 & 4): 13-18.

[PDF] 1963. Animal species, evolution and geographic isolation. Systematic Zoology 12(4): 202-204.

[PDF] 1964. The evolution of mating behavior in arthropods. In K. C. Highnam (ed.), Insect Reproduction. Symposium of the Royal Entomological Society of London 2: 78-94.

[PDF] 1964. Communicative systems of animals: Acoustic behavior. Science 114: 713-715 (Book Review).

[PDF] 1966. The evolution of cricket chirps. Natural History 75(9): 26-31.
[PDF] Reprinted in Niles Eldredge (ed.), The Natural History Reader in Evolution. New York: Columbia University Press, pp. 79-86 (1987).

1967. The beetle. Introductory Zoology Laboratory Exercise. Ann Arbor, Mich., Edwards Brothers Printers, 6 pp.

[PDF] 1967. Acoustical communication in arthropods. Annual Review of Entomology 12: 495-526.

[PDF] 1967. Seasonal and daily chirping cycles in northern spring and fall field crickets. Ohio Journal of Science 67(4): 200-209. (R.D. Alexander and G. H.. Meral).

[PDF] 1967. Cannibalism during copulation in the brown bush cricket, Hapithus agitator Uhler (Gryllidae). Florida Entomologist 50(2): 75-87. (R.D. Alexander and D. Otte).

[PDF] 1967. Basic research and public support. Science 157(3785): 134-136.

[PDF] 1967. Protests unexpected editorial changes. Science 157(3795): 1377.

[PDF] 1967. The evolution of genitalia and mating behavior in crickets (Gryllidae) and other Orthoptera. Miscellaneous Publications of the University of Michigan Museum of Zoology 133: 1-62. (R.D. Alexander and D. Otte).

[PDF] 1967. A critique of Animal Species and Evolution by Ernst Mayr, pp. 1-141 (Results of a seminar with eight graduate students).

[PDF] 1968. A comparative review (of On Aggression and The Territorial Imperative). BioScience 18(3): 245-248 (R.D. Alexander and D. W.. Tinkle).
[PDF] Excerpted in P. B. Weisz (ed.), The Contemporary Scene: Readings on Human Nature, Race, Behavior, Society, and Environment (New York: McGraw-Hill, 1970), pp.35-37.

[PDF] 1968. Life cycle origins, speciation, and related phenomena in crickets. Quarterly Review of Biology 43(1): 1-42.  [A “List of corrections and explanatory notes” for this paper is here.]

[PDF] 1968. Animal behavior and systematics. In: Proceedings of the Summer Institute in Systematics at the Smithsonian Institution, 25 June – 14 July 1967. Washington, DC: Smithsonian Institution, pp. 84-132.

[PDF] 1968. Arthropods. In: Sebeok, T. A., ed. Animal Communication: Techniques of Study and Results of Research. Bloomington IN: Indiana Univ. Press, p.167-216.

[PDF] 1969. Comparative animal behavior and systematics. In Proceedings of the International Conference on Systematics (Ann Arbor, Michigan, July 1967) National Academy of Sciences. Pub. 1962: 494-517.


[PDF] 1971. The search for an evolutionary philosophy of man. Proceedings of the Royal Society of Victoria, Melbourne 84: 99-120.

[PDF] 1972. Behavior associated with reproduction in field crickets. In A Laboratory Manual: Introduction to Biology, Campus Publishers, Shappirio, Frye, and Ray (eds.), Ann Arbor, Michigan.

1972. Diversity of organisms. In A Laboratory Manual: Introduction to Biology, Campus Publishers, Shappirio, Frye, and Ray (eds.), Ann Arbor, Michigan.

[PDF] 1972. The singing insects of Michigan. Great Lakes Entomologist 5(2): 33-69. (R.D. Alexander, A. E. Pace, and D. Otte).

[PDF] 1972. Saying it with sights and sounds. In Marvels of Animal Behavior. Washington, D.C.: National Geographic Society, pp. 88-111.

[PDF] 1973. The robust conehead: Two widespread sibling species (Orthoptera: Tettigoniidae: Neoconocephalus “robustus”). Ohio Journal of Science 73(6)321-330 (T. J. Walker, J. J. Whitesell, and R. D. Alexander).

[PDF] 1974. Evolution of sterile castes in the social insects: Kin selection or parental manipulation? Florida Entomologist 57(1): 32.

[PDF] 1974. The evolution of social behavior. Annual Review of Ecology and Systematics 5: 352-383.

[PDF] 1974. Jasper Loftus-Hills (Obituary), Copeia 3: 812.

[PDF] 1974. Creativity from a scientist’s viewpoint. Diagonalia: The University of Michigan LS&A College Newsletter (Dec. 1974).

[PDF] 1975. Natural enemies in place of poisons. Natural History 84(1): 92-95. (Book review of Paul DeBach’s Biological Control by Natural Enemies).

[PDF] 1975. The search for a general theory of behavior. Behavioral Science 20: 77-100.

[PDF] 1975. Natural selection and specialized chorusing behavior in acoustical insects. In: Insects, Science and Society, Proceedings of the Comstock Centennial Celebration, David Pimental (ed.), Cornell Univ., NY.: Academic Press, pp. 35-77.

[PDF] 1975. Insect control. Science 188: 206-208 (P. W.. Sherman and R.D. Alexander).

[PDF] 1977. Local mate competition and parental investment: patterns in social insects. Science 196: 494-500 (R.D. Alexander and P. W. Sherman).

[PDF] 1977. Natural selection and the analysis of human sociality. In Changing Scenes in the Natural Sciences, 1776-1976. C. E.. Goulden (ed.). Bicentennial Symposium Monograph, Philadelphia Academy of Natural Sciences, pp. 283-337.

[PDF] 1977. Evolution, human behavior, and determinism. Proceedings of the Biennial Meeting of the Philosophy of Science Association (1976), 2: 3-21.

[PDF] 1977. [Review of] The Use and Abuse of Biology by Marshall Sahlins. American Anthropologist 79: 917-920.

[PDF] 1978. Natural selection and societal laws. In The Foundations of Ethics and its Relationship to Science, Vol. 3. Morals, Science, and Society. Hastings-on-Hudson, NY, Feb. and June 1977. T. Englehardt, Jr. and D. Callahan (eds.), pp. 249-290. [Reprinted in The Roots of Ethics: Science, Religion, and Values. The Hastings Center, Plenum Press, NY, 1981. D. Callahan and T. Engelhardt, Jr., (eds.), pp. 265-306.]

[PDF] 1978. Rejoinder to Kenneth Schaffner. In The Foundations of Ethics and its Relationship to Science, Vol. 3. Morals, Science, and Society. Hastings-on-Hudson, NY, Feb. and June, 1977. T. Englehardt, Jr. and D. Callahan (eds.), pp. 304-308. [Schaffner’s chapter is here;  it is a response to RDA’s “Natural selection and societal laws”, above.]

[PDF] 1978. Letter to editor of Michigan Daily (Rejoinder to comment on rape). [In this letter (at the bottom right of the page, entitled “sociobiology”), RDA responded to a false comment about him in this article by the “Sociobiology Study Group, Ann Arbor Science for the People”. Their comment is at the top of the middle column of the article “The Sociobiology myth”. That group then responded to his letter here (letter entitled “no misquote”)].

[PDF] 1978. Evolution, creation, and biology teaching. American Biology Teacher 40: 91-104.

[PDF] 1978. The challenge of Darwinism. Rackham Reports (University of Michigan Graduate School) 4: 1-2. [Reprinted in Michigan Discussions in Anthropology 3: 144-148.]

[PDF] 1978. Biology, determinism, and human behavior: A response to Slobodkin, Michigan Discussions in Anthropology 3: 154-166. [The original article to which RDA is responding is here. Slobodkin then wrote a response here.]

[PDF] 1978. Group selection, altruism and the levels of organization of life. Annual Review of Ecology and Systematics 9: 449-474. (R.D. Alexander and G. Borgia).

[PDF] 1978. [Review of] The Collected Papers of Charles Darwin edited by Paul H. Barrett. Michigan Academician 9: 109-111.

[PDF] 1979. Natural selection and social exchange. In Social Exchange in Developing Relationships, R. L. Burgess and T. L. Huston (eds.), NY: Academic Press. pp. 197-221.

[PDF] 1979. The origin and basis of the male-female phenomenon. In Sexual Selection and Reproductive Competition in Insects, M. F. Blum and N. A.. Blum (eds.), Academic Press, pp. 417-440 (R.D. Alexander and G. Borgia).

[PDF] 1979. Evolution and culture. In Evolutionary Biology and Human Social Behavior: An Anthropological Perspective. North Scituate, Mass.: Duxbury Press, pp. 59-85, N. A. Chagnon and W. G. Irons (eds.).

[PDF] 1979. Sexual dimorphisms and breeding systems in pinnipeds, ungulates, and humans. In Evolutionary Biology and Human Social Behavior: An Anthropological Perspective., N.A. Chagnon and W. G. Irons (eds.), North Scituate, Mass.: Duxbury Press, , pp. 402-435 (R. D. Alexander, J. L. Hoogland, R. D. Howard, K. L. Noonan, and P. W. Sherman).

[PDF] 1979. Concealment of ovulation, parental care and human social evolution. In Evolutionary Biology and Human Social Behavior: An Anthropological Perspective, N. A. Chagnon and W. G. Irons (eds.), North Scituate, Mass.: Duxbury Press, pp. 436-453 (R. D. Alexander and K. M. Noonan).

[PDF] 1979. Sexuality and sociality in humans and other primates. In Human Sexuality: A Comparative and Developmental Perspective, H. Katchadourian (ed.). Berkeley, Calif., Univ. of Calif. Press. pp. 81-97. [This chapter can stand alone, but in part responds to the previous chapter, by Jane Lancaster, here.]

[PDF] 1979. Human sexuality and evolutionary models. In Human Sexuality: A Comparative and Developmental Perspective, H. Katchadourian (ed.). Berkeley, Calif, Univ. of Calif. Press, pp. 107-112. [This piece is sensible by itself, but is (rather oddly) billed as a commentary on the chapter (mostly by Frank Beach) of which it forms the final section. The chapter in its entirety, including RDA’s comments, is here.]

[PDF] 1979. Evolution, social behavior, and ethics. In The Foundation of Ethics and Its Relationship to Science,IV. Science, Morals, and Society, T.E. Engelhardt and D. Callahan (eds). Hastings-on-Hudson, New York: The Hastings Institute. pp. 124-155. [Reprinted in The Roots of Ethics: Science, Religion, and Values. The Hastings Center, Plenum Press, NY, 1981. D. Callahan and T. Engelhardt, Jr., (eds.), pp. 307-338.]

[PDF] 1979. Comment on Boucher, et al. Michigan Discussions in Anthropology 4: 34-35. [The original article to which RDA is responding is here].


[PDF] 1980. Field crickets (Gryllus and Acheta) of California and Baja, California (Orthoptera: Gryllidae: Gryllinae), Transactions of the American Entomological Society 106: 327-356 (D. B. Weissmann, D. F. C. Rentz, R. D. Alexander, and W. Loher).

[PDF] 1980. The Social Insects [Review of H.R. Hermann, ed., The Social Insects. New York: Academic Press]. Bioscience 31: 64-65 (R. D. Alexander and K. M. Noonan).

[PDF] 1981. Introduction to Natural Selection and Social Behavior: Recent Research and New Theory.  R. D. Alexander and D. W. Tinkle (eds), New York: Chiron Press, pp. xi-xii.

[PDF] 1981. Evolution, culture, and human behavior: some general considerations. In Natural Selection and Social Behavior: Recent Research and New Theory. R. D. Alexander and D. W. Tinkle (eds), New York: Chiron Press, pp. 509-520.

[PDF] 1982. Culture theory: the developing synthesis from biology, Human Ecology 10(3): 383-400, (M. Flinn and R. D. Alexander).

[PDF] 1982. Biology and the moral paradoxes. Journal of Social and Biological Structures 5: 389-395. [Reprinted, 1983, In Law, Biology, and Culture, M. Gruter and P. Bohannan (eds.), Santa Barbara, Calif.: Ross-Erikson, Inc., pp. 101-111].

[PDF] 1985. Genes, consciousness, and behavior theory. In A Century of Psychology as Science, Sigmund Koch and David E. Leary (eds.). N.Y.: McGraw-Hill, pp. 783-802.

[PDF] 1985. A biological interpretation of moral systems, Zygon 20: 3-20.

[PDF] 1986. Ostracism and indirect reciprocity: the reproductive significance of humor. In Ethology and Sociobiology 7: 253-270R, Masters and M. Gruter (eds).
[PDF] Translated and republished in German as: Ostrazismus und Indirekte Reziprozität: Die reproduktive Bedeutung des Humors. pp.79-99 in Gruter, Margaret & Rehbinder, Manfred (eds.): Ablehnung – Meidung – Ausschluß: multidisziplinäre Untersucungen über die Kehrseite der Vergemeinschaftung. Berlin (Duncker und Humblot), 1986.]

[PDF] 1986. Biology and law. In Ethology and Sociobiology 7: 167-173R. Masters and M. Gruter (eds.).
[PDF] Translated and republished in German as: Recht, Biologie und Sozialverhalten. pp. 29-36 in: Gruter, Margaret & Rehbinder, Manfred (eds.): Ablehnung – Meidung – Ausschluß: multidisziplinäre Untersucungen über die Kehrseite der Vergemeinschaftung. Berlin (Duncker und Humblot), 1986.

[PDF] 1987. Human hips, breasts, and buttocks: Is fat deceptive? Ethology and Sociobiology 8(4): 249-257. B. S. Low, R. D. Alexander, and K. M. Noonan.

[PDF] 1987. The real origin of the species, Washington Post (June 28, p. C3). Invited editorial.

[PDF] 1987. The crickets of New Caledonia (Gryllidae). Proceedings of the National Academy of Sciences, Philadelphia 139: 375-457. (Daniel Otte, R.D. Alexander, and W. Cade).

[PDF] 1988. The evolutionary approach to human behavior: What does the future hold? In Human Reproductive Behavior: A Darwinian Perspective, London: Cambridge Univ. Press, L. L. Betzig, M. Borgerhoff Mulder, and P. W. Turke (eds.), pp. 317-341.

[PDF] 1988. Über die Interessen der Menschen und die Evolution von Lebensabläufen. In Die Herausforderung der Evolutionsbiologie, H. Meier (ed). Munich: R. Piper GmbH & Co. KG, pp. 129-171. Translated by Agnes von Cranach.

[PDF] 1988. Knowledge, intent and morality in Darwin’s world. [A review of Robert J. Richards, Darwin and the Emergence of Evolutionary Theories of Mind and Behavior (Chicago: University of Chicago Press, 1987)]. Quarterly Review of Biology 63:441-443.

[PDF] 1988. Response to Judith Anderson’s comments on Low, Alexander, and Noonan (1987), Ethology and Sociobiology 9: 325-328. (B. S. Low, R. D. Alexander, and K. M. Noonan).

[PDF] 1989. The evolution of the human psyche. In The Human Revolution, C. Stringer and P. Mellars (eds), Univ. of Edinburgh Press, pp. 455-513.

[PDF] 1989. The evolutionary significance of the aesthetic sense, LSA Magazine, Spring, Volume 12, Number 2, pp. 22-24.

[PDF] 1989. T. H. Hubbell, 1898-1989 (Obituary). Metaleptea: The Orthopterists’ Newsletter 11(1-2):3-5. [The entire issue is included in this PDF]


[PDF] 1990. Epigenetic rules and Darwinian algorithms: The adaptive study of learning and development, Ethology and Sociobiology 11: 241-303.

[PDF] 1990. How did humans evolve? Reflections on the uniquely unique species. Special Publication of the University of Michigan Museum of Zoology 1: 1-38.

[PDF] 1991. The evolution of eusociality. In The Biology of the Naked Mole Rat, P. W. Sherman, J. Jarvis, and R. D. Alexander (eds). Princeton Univ. Press. (R. D. Alexander, K. M. Noonan, and B. Crespi).

[PDF] 1991. Unanswered questions about naked mole rats. In The Biology of the Naked Mole Rat, Princeton Univ. Press. P. W. Sherman, J. Jarvis, and R. D. Alexander (eds).

[PDF] 1991. An ethogram of the naked mole-rat. In The Biology of the Naked Mole-Rat, Princeton Univ. Press (E. Lacey, J. Pepper, P. W. Sherman, and R. D. Alexander).

[PDF] 1991. A Review of the Genus Gryllus (Orthoptera: Gryllidae), with a new wpecies From Korea, Great Lakes Entomologist, Vol. 24, No. 2, pp. 79-84.

[PDF] 1991. Social learning and kin recognition: an addendum and reply to Sherman, Ethology and Sociobiology 12: 387-399.

[PDF] 1993. Biological considerations in the analysis of morality. In Evolutionary Ethics, M. H. Nitecki (ed), State University of New York Press.

[PDF] 1993. A biologist’s approach to human nature. In Man and Creation, M. Bauman, L. Roche, and J. Corombos (eds). Hillsdale, MI, Hillsdale College Press, pp. 191-207.

[PDF] 1994. A report from the director. The University of Michigan Museum of Zoology, October 1994. Submitted to Edie N. Goldenberg, Dean, College of Literature, Science & The Arts. 69 pp.

[PDF] 1995. The view from the president’s window: Impressions from the Santa Barbara Meeting. Human Behavior and Evolution Newsletter, Volume 4(2), September 1995.

[PDF] 1996. The view from the president’s window. Human Behavior and Evolution Newsletter. Volume 5(1), February 1996.

[PDF] 1996. The view from the president’s window: Dilemmas of Darwinists: A few tips of the iceberg. Human Behavior and Evolution Newsletter, Volume 5(2), Fall 1996.

[PDF] 1996. New Records of Michigan Cicadidae (Homoptera), with notes on the use of songs to monitor range changes. Great Lakes Entomologist. Vol. 29(3), pp. 165-169. (D.C. Marshall, J. R. Cooley, R. D. Alexander, and T. E. Moore.)

[PDF] 1997. The view from the president’s window: On God, and such. Human Behavior and Evolution Newsletter, Volume 6(1), Spring 1997.

[PDF] 1997. Special report to Dean Edie N. Goldenberg. The University of Michigan Museum Of Zoology, 4 June 1997, ix + 42 pp.

[PDF] 1997. Evolutionary perspectives on insect mating. In: The Evolution of Mating Systems in Insects and Arachnids. J. C. Choe and B. J. Crespi (eds.) Princeton University Press, pp. 4-31. (Richard D. Alexander, David Marshall, and John Cooley).


[PDF] 2000. William D. Hamilton remembered. Natural History, June 2000, Vol. 109, Issue 5.

[PDF] 2001. Range bulls. Transactions of the American Entomological Society 127(3): 289-290.

[PDF] 2003. Crickets. In: Encyclopedia of Insects. Vincent H. Resh and Ring T. Cardé (eds.) New York, Academic Press, pp. 265-269 (Richard D. Alexander and Daniel Otte).

[PDF] 2005. Evolutionary selection and the nature of humanity. In V. Hosle and Ch. Illies (eds). Darwinism and Philosophy (Notre Dame: University of Notre Dame Press), pp. 424-495.

[PDF] 2005. The ills of society can influence “killing gene”. Ann Arbor News. [This is a reply to David Buss’ book The Murderer Next Door: Why the Mind is Designed to Kill].

[PDF] 2006. The challenge of human social behavior: A review essay stimulated by Hammerstein, Peter (Ed.) 2003. Genetic and Cultural Evolution of Cooperation. (Cambridge, Massachusetts, and London, England: MIT Press). Evolutionary Psychology 4: 1-32.

[PDF] 2007. Runaway social selection in human evolution. In Gangestad, Steven W. and Jeffry A. Simpson (eds.), The Evolution of Mind: Fundamental Questions and Controversies. New York: Guildford Press, pp. 249-255. (Mark Flinn and R. D. Alexander).

[PDF] 2008. Evolution and human society. A lecture delivered at the June 2008 Kyoto, Japan, meeting of the Human Behavior and Evolution Society. Human Behavior and Evolution Newsletter, Summer 2008, pp. 5-15; with supplements. [This was RDA’s keynote address upon receiving the HBES Distinguished Scientific Award for Lifetime Contribution to Human Behavior and Evolution. This expanded version, finalized on 26 December 2009, contains (1) an abstract or introduction to the lecture; (2) the lecture itself, entitled “Understanding the human species and its immediate ancestors”; (3) the associated handout, entitled “A brief recounting of recent evolutionary findings”; (4) an appendix of explanatory footnotes; and (5) a table comparing relevant traits among the apes including humans. For the original HBES newsletter version, click here.]

[PDF] 2008. Let’s examine both sides of the coin of human understanding. Ann Arbor News: Other Voices, 17 July. A scan of the original paper copy is here. [This is a reply to a 12 July letter by Ronald G. Larson entitled “Evolutionary theory doesn’t explain it all”].

[PDF] 2008. Natural complexity is commonplace. Ann Arbor News: Opinion, 10 August. [A reply to a 4 August letter by Lahki Goenka].

[PDF] 2009. An HBES hero: A special tribute to Dr. Billy E. Frye. Human Behavior and Evolution Society Newsletter. Winter 2009, 2 pp. (Richard D. Alexander, William G. Irons, and Napoleon E. Chagnon).

[PDF] 2010. Understanding ourselves. In L. C. Drickamer and D. A. Dewsbury (eds.), Leaders in Animal Behavior: The Second Generation. New York: Cambridge University Press, pp. 1-37. [This paper, as with the others in this two-volume set, is a professional autobiography.]

[PDF] 2013. Religion, evolution and the quest for global harmony. In K. Summers and B. Crespi (eds.), Human Social Evolution: The Foundational Works of Richard D. Alexander (New York: Oxford University Press), pp.384-425. [Included in this file is a poem by RDA selected by the editors, and an introduction by William Irons]

[PDF] 2013. The distribution of periodical cicada Brood III in 1997, with special emphasis on Illinois (Hemiptera: Magicicada spp.). American Entomologist 59: 9-14. (Cooley, J.R., Marshall, D.C., Richards, A.F., Alexander, R.D., Irwin, M.D., Coelho, J.R. and Simon, C.).

[PDF] 2014. Darwin’s challenges and the future of human society. In Predicting the Future in Science, Economics, and Politics, F. Wayman, P. Williamson, and B. Bueno de Mesquita (eds.), Ann Arbor: University of Michigan Press, pp. 55-108.

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Unpublished Professional Manuscripts

[PDF] 1988. Evolution of the human psyche and some of its special activities (music emphasized here). Unpublished figure, 10 April.

[PDF] 1992. A theory of music. Unpublished ms, 11 December.

[PDF] 1993. Learning. Unpublished ms, 31 October.

1997. Understanding Humanity: The Human Species in Evolutionary Perspective. Prepared as a text for Biology 494, University of Michigan (assisted by Andrew F. Richards).

1998. Reply to Matt Cartmill. Unpublished ms, 16 March.

[PDF] 1998. Using evolutionary selection to help understand humanity. Unpublished ms, 16 September.

1999. The Williamsburg Address. Unpublished ms, 11 January.

[PDF] 1999. The three great gaps in biology. Unpublished ms, 8 February.

1999. Comment on Ewald. 15 February.

[PDF] 1999. Trying to make evolutionary sense of some human uniqueness. Unpublished handout to the University of Michigan Evolution and Human Adaptation Program, March.

1999. Some contrasts in evolutionary routes to social systems and social brains. Unpublished ms, 9 March.

2000. On the advocacy of intelligent design of life. Unpublished ms.

2001. Complexity and adaptation. Unpublished ms, 31 October.

[PDF] 2002. The woes of the world. Unpublished ms, 4 February.

[PDF] 2002. The biology of pleiotropies. Unpublished ms, 15 April.

[PDF] 2003. Who is misusing the moral module: the self-declared godless, the faith-based, or both? Unpublished ms, 10 April.

2003. Forms of evolutionary selection. Unpublished ms, July.

[PDF] 2004. The nature of human morality. Unpublished ms.

[PDF] 2004. Prayer for the kindred of the world. Unpublished ms.

[PDF] 2006. The concept of God and the meaning of life. Unpublished ms. [Included as an addendum is a publication-friendly version of the “Prayer for the kindred of the world” (c.f. 2004 above).]

[PDF] 2006. Review of The Roots of Morality by Maxine Sheets-Johnstone. Unpublished ms, 16 May.

[PDF] 2007. Bragging rights: the sides of the social cooperation coin. Unpublished ms, 23 February.

[PDF] 2007. Have God and evolution constructed humans jointly? Unpublished ms, 1 May.  [A version from 23 February 2007 is here.]

[PDF] 2007. Self-interest and its alternatives. Unpublished ms, 12 March.

[PDF] 2007. Responses to the online comments [an Amazon.com review] of the economist Herbert Gintis regarding The Biology of Moral Systems. 14 April.

[PDF] 2008. On ultimacy in the life sciences. Unpublished ms. [a 2004 version is here].

2011. The mockingbird, or, Religion: an evolved adaptation? And what if it is?! Unpublished ms, 28 May.

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Nonprofessional Works

1969. Excerpts from an Australian Journal. xiv + 495 pp. [Unpublished, distributed privately.]

1975. Pop: 111 Stories about My Father. 139 pp., 19 drawings. [Unpublished, distributed privately.]

1977. The Clarksburg Cat, and Other Things That Happened along the Way: 41 Stories and Verses, 141 pp. [Unpublished, distributed privately.]

1979. A Part of the Hunting: 66 Verses written between 1949 and 1979. vii + 102 pp. [Unpublished, distributed privately.]

[PDF] 1988. Moving my family to Michigan, 9 pp. [Unpublished, originally intended for Notes on the Faculty; originally written in August 1988, revised in the 2010s].

[PDF] 1991. Mom’s Story. The Life of Katharine Elizabeth Heath Alexander Stutzenstein (edited).128 pp., 101 photographs, 2 maps, 1 drawing. Manchester, MI: Woodlane Farm Books [compiled and edited by RDA– text is mostly from Katharine’s perspective, originating in interviews by RDA]. In 1991 and 1992, RDA created two lists of minor errata: here.

1997. Playin’ Cowboy: 71 Verses, Songs, Stories, and Sketches, 160 pp. Typescript, distributed privately. Republished in 2006 as Playin’ Cowboy: The Coontail Blue and Other Horse Tales. 321 pp. Manchester, MI: Woodlane Farm Books.

1998. Dick Alexander’s Song Book I. 500+ Songs I Have Known and Sung: 1948-1998. 556 pp. [Unpublished, distributed privately.]

1998. Dick Alexander’s Song Book. II. 200+ Songs from Old and Out-of-Print Books. 426 pp. [Unpublished, distributed privately.]

1998. Dick Alexander’s Song Book III. 88 Songs and Verses I Have Written, Modified, or Discovered, and Saved. 1951-1998. 258 pp. [Unpublished, distributed privately.]

1998. The Harley Davidson reining horse. Poem and drawing. National Reining Horse Association Reiner, February 1996, p. 115.

[PDF] 2000. An old-time vet. Manchester Enterprise, July 6, pp. 4,10.

2001. Letter to the editor on imprinting of foals. The Horse. 

[PDF] 2001. Club 48: A Personal Account of Blackburn College and the Men of Butler Basement. 1946-48. Manchester, MI: Woodlane Farm Books, 128 pp., 51 photographs.

2001. Teaching Yourself to Train Your Horse. Simplicity, Consistency, and Common Sense from Foal to Comfortable Riding Horse. Manchester, MI: Woodlane Farm Books, xiv + 276 pp., 334 color photographs, 3 color diagrams, three appendices, bibliography.
[In 2002 RDA posted a list of (minor) errata: here.]

Essays on Horses and Horse Training:
[This informal series was originally posted on the (now defunct) Woodlane Farm website; each essay was conceived as a supplement to his 2001 book Teaching Yourself to Train Your Horse.]

[PDF] 2001. Keeping the young horse’s trust while riding.
[PDF] [Reprinted in a Finnish translation in Western Riders Finland, v.2/2002, transl. Juha Leppänen, pp. 26-27. This PDF contains the entire issue].

[PDF] 2002. Understanding rewards and punishment in horse training.

[PDF] 2002. Targeting, clicker-bridging, and positive and negative aspects of horse training. (Richard D. Alexander and Cynthia Kagarise Sherman).

[PDF] 2002. Four ways of training (Richard D. Alexander and Cynthia Kagarise Sherman). [Includes an introduction to Cindy].

[PDF] 2002. Two seconds a day can gentle your horse.

[PDF] 2002. Holding on with upper legs.

[PDF] 2002. Long latigos.

[PDF] 2002. On using ropes to lift a young horse’s legs and questions about secure tying.

[PDF] 2002. An accidentally omitted reference on advanced riding.

2003. The Red Fox and Johnny Valentine’s Blue-Speckled Hound. ca. 93 pp., 21 watercolors by John Megahan, numerous pencil sketches of red foxes by William Berry. Woodlane Farm Press.

2003. The old German horse farmer of Freedom Township. Poem. Manchester Enterprise, Manchester, MI. Thursday 12 April 2003, p.9A.

[PDF] 2005. Pop’s Story: A Midwestern Farm Boy’s Memories of Times with His Father. 162 pp. Manchester, MI: Woodlane Farm Books.

[PDF] 2006. Looking back: Ann Arbor balers. Manchester Enterprise, Manchester, MI. Thursday 12 October. p.2B.

2006. Looking back: The last threshing run in Piatt County, Illinois. Manchester Enterprise, Manchester, MI. Thursday 28 September. p.3B. Photographs by Mark O’Brien.

2007? Op Ed on evolution and creationism. Ann Arbor News.

2008. Reply to __ re creationism and evolution. Ann Arbor News 2008.

2009. Thumping On Trees. 43 pp. 3 stories. Illustrations by John Megahan. Manchester, MI: Woodlane Farm Books.

[PDF] 2012, unfinished. Special Babies & Their Parents: Calves, Colts, Kids, and Others. [This is a portion of a proposed book, containing examples of true observations of animal babies by RDA and Megan, with an addendum about the distinctiveness of the human baby.]

[PDF] 2015. Notes On The Faculty– Including, of Course, a Few Curious Examples. ix+174 pp. Manchester, MI: Woodlane Farm Books. This volume includes the following works:

      • Introduction
      • The ancient profs who never die (poem)
      • Theodore Huntington Hubbell: Long-Time Director of the University of Michigan Museum of Zoology
          • Pure altruism? (cartoon)
      • Donald J. Borror: Lessons I Learned as His First Doctoral Student. [This was previously distributed as “My interactions with Donald J. Borror, as his first doctoral student”, from 9 April 2003.]
          • Donald J. Borror, Odonata, Taxonomy, Bird Song (cartoon)
      • Josef N. Knull: Curator’s Curator
          • Josef Knull, Buprestidae, Taxonomy, Curator of the Museum (cartoon)
      • Edward S. Thomas: Ohio Naturalist and Honorary Doctorate
      • Donald W. Tinkle: Outstanding Lecturer, Ecologist, and Museum Director
      • Carl Venard
          • Carl Venard, Medical Entomology (cartoon)
      • Ralph Davidson
          • Ralph Davidson, Economic Entomology (cartoon)
      • Frank Fisk
          • Frank Fisk, Physiological Entomology (cartoon)
      • Alvah Peterson
          • Alvah Peterson, Immature Insects, Biological Control (cartoon)
      • Dwight M. Delong
          • Dwight M. Delong, Homoptera, Ecology (cartoon)
      • Jasper Loftus-Hills: Australian Biologist
      • Carl Walter Campbell: A Sixty-Year Friendship. An Essay Dedicated to the Campbell Family [previously distributed in a slightly different form as “My Sixty-Year Friendship with Carl Walter Campbell”, from 13 January 2007.]
      • William D. Hamilton: Remembered. [The first section of this essay was previously published in Natural History in 2000 (see above).]
          • Reverse Reciprocity (poem)
      • Billy E. Frye: A Special Tribute (with William G. Irons and Napoleon A. Chagnon). [The first section of this part was previously published in the HBES Newsletter as “An HBES Hero: A Special Tribute to Billy E. Frye” in 2009 (see above).]
          • University Administrations (poem)
      • Kybe Hartweg: Curator of Reptiles (poem and drawing)
      • Charles Walker: Professor of Frogs’ Lives (poem)
      • Claude Hibbard: Hibbie- Fossil Man (poem)
      • Aubrey Hart: Museum Shop Man (poem)
      • Noble Porter Heath II: My Mother’s Father (essay and drawing)
      • A Place for Sam (poem and drawing)
      • Henry P. Eames: An Old-Time Veterinarian [This was previously published in the Manchester Enterprise, Manchester, Michigan, on July 6, 2000, as “An old-time vet” (see above).
      • Dustin Stinson: Veterinarian
          • The Song and Story of The Clarksburg Cat (poem)
      • Donald R. Meyer
      • John Wolfe: Hidden Roots and the Limits of Consciousness
          • Big Bluestem (poem)
      • The Biology of Mole Systems (true story)
      • On Getting Things Straight (true story)
      • New Museum Folk Song (poem)
      • Remarks for the Illinois State University Commencement (true story)
      • On Wisdom (essay)
      • Horse Training and Searching for the Truth (essay)
      • [dog] (drawing)
      • After decades rife with science strife… (poem)
      • [cricket on leaf] (drawing)
      • [roping a steer from a distance] [drawing, silhouette]
      • It is surely time for the adaptive structures… (poem)
      • [probable self-portrait on horseback] (drawing)

[PDF] 2016. White Man’s Fire: Tales Of Two Cricket Hunters in Australia, 1968-1969. 209 pp. Illustrations and photographs by Daniel Otte. Manchester, MI: Woodlane Farm Books.

Of ladybugs and their betters. Poem. The Wanderer.

Letter to Ann Arbor News re Bridgewater Bank Tavern (quoted).

Stealing Watermelons: Tales from Sangamon Township, ~300 pp. Manchester, MI: Woodlane Farm Books.

The Mockingbird’s River Song. Poems, songs, essays, and stories. ~300 pp. Manchester, MI: Woodlane Farm Books.

Letter to columnist in Quarter Horse News re horse bridle bits (included in the column).

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This website was launched in June 2010 and is a work in progress.
For more information or to become involved,
contact David Lahti at dclahti@gmail.com.

In addition to RDA of course, thanks to Lorrie Alexander, John Cooley, Daniel Otte, Eckart Voland, Mary Jane West-Eberhard, and especially Nancy Alexander for providing a few hard-to-find writings. Thanks to Michelle Mordukhaev for contributing to the scanning and posting. Thanks to Nancy Alexander and Lorrie Alexander for guidance and permission.

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